Thomas Hunt Morgan (1866-1945) proposes that Mendelian factors are arranged in a line on chromosomes.
Editors. "Birth of genetics: genes are carried on chromosomes and are the mechanical basis of heredity". The Literary Encyclopedia. First published 01 May 2009
[http://www.litencyc.com/php/stopics.php?rec=true&UID=6427, accessed 22 November 2014.]